Apple Teacher
Biomedical Research CITI Program Certification
Ph.D., M.S., University of Rochester
B.S., Pennsylvania State University
Professor Millen is a biologist whose work bridges the science of aging with student-centered teaching and mentorship. His academic path began with a fascination for biology at Penn State, followed by graduate and postdoctoral training at the University of Rochester, where he studied cellular processes in yeast and their connection to aging and disease. These experiences solidified his passion for research and a growing enthusiasm for teaching, ultimately guiding him to St. John Fisher University.
His teaching is built on the belief that students learn best through active participation, reflection, and real-world application. He designs courses that invite students to ask questions, experiment, and connect scientific ideas to meaningful contexts. Signature experiences such as The Longevity Games place students in partnership with older adults, blending biology with community engagement. Whether in foundational courses or interdisciplinary seminars, he strives to create classrooms that are rigorous yet welcoming, where curiosity and collaboration drive learning.
His scholarship focuses on aging biology, small-molecule therapeutics, and the integration of artificial intelligence into research and education. In the lab, he investigates compounds that may extend lifespan or protect against the side effects of cancer treatments. Alongside this molecular work, he develops AI-based tools that enhance both drug discovery and student learning. These dual tracks reflect his commitment to scientific discovery while mentoring the next generation of biologists. At Fisher, he sees teaching, research, and service as deeply interconnected. His is to continue building opportunities where students can contribute to real science, gain confidence in their abilities, and prepare to carry their curiosity forward into their careers and communities.
Selected Publications
- Using generative AI for interview simulations to enhance student research skills in biology education, J Microbiol Biol Educ 26(2), 2025.
- Positive cardioprotective effects of topoisomerase II poison, LS1, in doxorubicin treatment, Biol Biomed Sci 96(2):59–62, 2024. Co-authored with K. Keenehan.
- Lifespan-extending properties of Kalanchoe daigremontiana plant extracts in Saccharomyces cerevisiae, Biol Biomed Sci 93(1):12–21, 2022. Co-authored with O. Marziale, X. Somoulay, S. Anand, MAB Herman, and J. Mattiaccio.
- Checkpoint regulation of DNA topoisomerase II by phosphorylation and SUMOylation, Nucleic Acids Res 45(10):5269–5283, 2017. Co-authored with G. Tombline, B. Polevoda, M. Rapaport, B. Baxter, K. Wennerberg, J. Madrey, M. Van Meter, J. Nitiss.